ZooKeys is a peer-reviewed, open-access, rapidly produced journal launched to support free exchange of ideas and information in systematic zoology. All papers published in ZooKeys can be freely copied, downloaded, printed and distributed at no charge for the reader. Authors may retain all other rights on their works. Authors are thus encouraged to post the pdf files of published papers on their homepages or elsewhere to expedite distribution. Papers are published both online and in the traditional printed format, in full compliance with the current requirements of ICZN.